The Middle Palaeolithic of Arabia: implications for modern human origins, behaviour and dispersals. (Research).
Petraglia, Michael D. ; Alsharekh, Abdullah
Introduction
The Middle Palaeolithic record has a central role for understanding
biocultural evolution and the origins of modern humans (e.g. Mellars
& Stringer 1989; Trinkaus 1989). Recent palaeoanthropological
syntheses indicate that the Middle Stone Age (MSA) of Africa may
coincide with a variety of key evolutionary events, including the
emergence of modern human morphology, the development of more complex
adaptations, and the dispersal of modern humans outside of Africa (e.g.
Foley & Lahr 1997; McBrearty & Brooks 2000). These syntheses
propose that H. helmei and H. sapiens produced MSA assemblages in
Africa, signalling a behavioural shift by c. 300-250 000 years ago. In
West Asia, Levantine Mousterian assemblages are associated with Homo
neanderthalensis and Homo sapiens, taken as evidence that human
populations moved through the Levantine corridor during episodes of
climatic deterioration and ameliorations (Bar-Yosef 1994). Various
researchers have examined the timing and routes of modern human
dispersals outside of Africa by examining patterns of morphological,
genetic, and archaeological diversity (e.g. Lahr & Foley 1994, 1998;
Quintana-Murci et al. 1999; Stringer 2000).
Despite its importance to understanding the emergence of modern
humans and the evolution of behaviour, the Middle Palaeolithic record of
the Arabian peninsula has been sidelined in palaeoanthropological
syntheses, and this marginalisation of the Arabian record prevails to
the present day despite its potentially crucial geographic position. At
least three Out of Africa dispersal routes may be envisioned which have
different evolutionary and behavioural implications. One is a northern
route through north-east Africa via the Sinai peninsula towards the
Levant, with subsequent movements into Arabia; a second is through the
Horn of Africa across the Bab al Mandab Strait, spreading along
coastlines and into inland zones; and a third is a coastal migration
along the Arabian Sea margin towards southern and south-eastern Asia,
some populations perhaps employing basic seafaring technology.
Although archaeological investigations on the Arabian Peninsula
have lagged behind those of surrounding regions, preliminary surveys
executed during the past twenty-five years have, in fact, resulted in
the identification of a number of Palaeolithic sites (Figure 1). While
serious problems remain in Arabian archaeology today, an aim of this
review is to bring the Middle Palaeolithic record of the region into the
fold of palaeoanthropological models, thereby providing for a more
holistic treatment of biocultural and behavioural patterns across the
Old World. As a consequence, the record of human occupation in the
Arabian Peninsula assumes great importance in discussions about hominin
biogeography, the pattern and route of hominin dispersals, and the use
of coastal zones for movement.
[FIGURE 1 OMITTED]
Geography and environments
The Arabian Peninsula is demarcated by the modern-day countries of
Saudi Arabia, Kuwait, Bahrain, Qatar, United Arab Emirates, Oman, and
Yemen. The landmass is large, measuring 2.3 million [km.sup.2] in total
extent. The peninsula is bounded on three sides by the Red Sea, the
Arabian Sea and the Arabian Gulf. In contrast, the northern area is a
vast open steppe that unrolls towards the Mediterranean with no major
geographic obstructions, likely leading to a free exchange of hominin
populations across a wide area.
In north-cast Africa, fluctuations in arid and humid phases have
been documented, Middle Palaeolithic occupations corresponding to lake
and spring deposits dating to between 175 000 and 70 000 BP (e.g.
Wendorf et al. 1993; Van Peer 1998; McBrearty & Brooks 2000).
Likewise, palaeoenvironmental scientists have observed marked changes in
the Arabian palaeoenvironments. A pollen core from the Arabian Sea,
ranging from the Holocene to more than 128 000 yrs, indicated that
glacial periods produced low sea levels and saline littoral and arid and
steppe inland conditions, while high sea-level corresponded with
savannah-type vegetation (Van Campo et al. 1982). Terrestrial
sedimentary deposits of the Arabian Peninsula were interpreted to
reflect changes in climate, glaciation events, and prevailing wind
systems (Glennie & Singhvi 2002). Arabia contains a diversity of
geomorphic features and sedimentary deposits, ranging from alluvial fans
to inland and coastal dunes and salt flats (Allison 1997). Moist
intervals are indicated by the formation of lakes, travertines with
fossil vegetation, and aggradation and resorting of deposits by local
stream flows. Arid phases occurred throughout the course of the late
Pliocene, Pleistocene, and Holocene, today resulting in the formation of
the vast desert known as the Empty Quartet (Rub' al-Khali).
Controlled by global climatic and atmospheric conditions, the large
inland depressions of the Rub' al-Khali in the south and the Nafud
Desert in the north may have undergone a similar pattern of infilling.
Analysis of the Lake Mundafan locality in the Rub' al-Khali showed
deep lacustrine deposits underneath the windblown sands (McClure 1976,
1978). A total of 28 radiocarbon dates in upper levels indicated at
least two episodes corresponding to periods of high water table, the
earlier phase ranging from 36 000 to 17 000 BP. Palaeolithic occurrences
associated with fluctuating lake and riverine conditions and aeolian deposits indicates that hominin populations had to cope with substantial
environmental change.
Fossilised coralline plains are located along the Red Sea coast of
Arabia. Palaeoclimatologists have related elevated coral reefs to
interglacial phases and submerged fossil corals and tributaries to
lowered sea levels during glacial periods (e.g. Lambeck & Chappell
2001; Lambeck et al. 2002). Examination of sea level fluctuations of the
Red Sea indicate changes in water depth and distance between Africa and
Arabia (e.g. Chapman 1971; Rohling 1994). Based on study of planktonic foraminifera, shallow sea levels are indicated in the Red Sea during
Oxygen Isotope Stage 6, c. 135 000 B.P (Rohling et al 1998), increasing
the likelihood for passage of hominins across the Bab al Mandab Strait,
either through an overland route or across a much reduced waterway.
Current knowledge and problems in the Arabian Middle Palaeolithic
A number of methodological, environmental, and financial factors
have been largely responsible for the slow growth of Palaeolithic
studies in Arabia. The severe arid settings in which many sites occur
has seriously hampered efforts to conduct Palaeolithic field studies in
depth. In addition, most archaeological research in Arabia tends to
focus on the study of historical periods, and concerted field efforts
with interdisciplinary scientific teams are not undertaken.
Middle Palaeolithic industries of the Arabian Peninsula have
traditionally been labelled as 'Mousterian' by most
investigators. Mousterian industries were distinguished from Lower and
Upper Palaeolithic assemblages based upon the relative frequency of
technological and typological attributes (i.e. relative frequency of
large bifaces, prepared cores, and blades). As is the problem in other
regions, industrial designations are arbitrary, consisting of relative
artefact proportions rather than absolute categories. Moreover, detailed
technological studies and quantification of tool types and reduction
techniques have not been performed, seriously inhibiting an
understanding of temporal and spatial characteristics. Although the term
Mousterian has been applied to the Arabian assemblages, questions remain
about the degree to which these assemblages share temporal-spatial
parallels with Levallois and discoidal industries in Europe, West Asia,
and North Africa. For this reason, the term 'Mousterian' is
reserved here to describe previously identified assemblages, whereas the
term 'Middle Palaeolithic' is used here as a cultural stage
without implying the presence of particular regional and stylistic
features.
Exacerbating technological problems, few reliable age estimates
have been recovered in Arabia. The poverty of chronological information
is not, however, due to a dearth of localities that can potentially
supply such information. Radiometric dating holds promise as appropriate
environments exist for such studies, including associations between
sites and deposits (e.g. volcanics, fossil corals, sediments). Caves,
buried sites, and variations in basin wide deposits with Palaeolithic
assemblages have been reported, although these have not been described
in detail (e.g. Garrard et al. 1981; Zarins et al. 1981; Amirkhanov
1994). Seemingly well-preserved, and buried sites have been identified
in alluvial situations and surveys indicate a high probability of
finding buried sites near lakeshores.
Of the identified sites, differences exist in their level of
observational recording by archaeologists, surface distributions often
recorded in a qualitative fashion (e.g. "widely dispersed, light
density artefact scatters", "highly clustered",
"dense sites"). Although researchers working on Palaeolithic
sites in West Asia have become aware of the role and influence of
formation processes in creating patterning (e.g. Goren-lnbar et al 1992;
Bar-Yosef 1993; Barton & Clark 1993), researchers in Arabia have not
paid attention to natural and cultural processes, leading to uncertainty
about the degree to which artefact distributions reflect hominin
activities. Additionally, artefact recovery on desert floors present
problems in understanding tool function, as edges have been modified and
rounded by chemical weathering and wind abrasion.
On the whole, survey results have indicated that Middle
Palaeolithic sites were more numerous in comparison to Acheulean and
Upper Palaeolithic localities. However, the recovery of this larger
sample of Middle Palaeolithic sites may be due to a number of
conditions, including biases in site visibility, differences in
preservation conditions, and mis-identification of later sites (presumed
to be only blade-bearing industries), or it may be related to functional
variations, such as occupation intensity and population size. Middle
Palaeolithic sites are exclusively known from Saudi Arabia and Yemen,
and large areas of the peninsula remain devoid of sites. The Gulf
States, including Oman, Bahrain, Kuwait, Qatar, and the United Arab
Emirates, either have no reported sites, or low numbers of temporally
problematic sites. The lack of sites in the Gulf States may reflect
variable occupation intensity in the peninsula, but it is also probable
that sites are submerged along littoral zones, or buried under large
sand tracts. The lack of sites in these areas may change if these
further surveys are initiated.
An evaluation of the Arabian Middle Palaeolithic
Environmental records
Middle Palaeolithic sites were located in different ecological
settings, including inland basins, coastal margins, and mountainous
zones. Site locations ranged widely, and included sand dunes, rivers of
different order, elevated areas and slopes overlooking valleys, lava
fields, and littoral margins. The presence of sites in the interior of
the Arabian Peninsula indicates that favourable conditions and resources
must have been present. Sites are often noted to be associated with
water sources, including rivers, streams, and lakes (e.g. Parr et al.
1978; McClure 1976, 1978; Zarins et al. 1980, 1981).
A particularly important set of sites was located along the Red Sea
coastal plain and lava fields (Zarins et al. 1980, 1981) (Figure 2). At
several localities, Levallois flakes, blades, cores, and other tool
types produced from lava were recovered. Artefacts were situated at +2m
above mean sea level and c. 75m from the sea, and in at least one case
the tools were firmly embedded in a coralline beach terrace. Little
other information is available about these sites, but their
identification is of extreme importance in identifying coastal
dispersals and potential subsistence practices of Middle Palaeolithic
hominins, as recently argued for African sites located on the Red Sea
(Walter et al 2000).
[FIGURE 2 OMITTED]
The Jubbah Basin sequence in the Great Nafud Desert of northern
Saudi Arabia provides important information about changes in environment
as seven major sedimentary units were identified in 26 metres of
deposits (Garrard et al 1981) (Figure 3). Lacustrine conditions were
thought to be represented in at least two periods of the Quaternary and
Holocene. The earliest level, Deposit 6, was a clay, indicating
accumulation under stillwater conditions over a long time interval.
Deposit 5b was an evaporite, suggesting increased aridity. This was
succeeded by Deposit 5a, a diatomite dated to 25 630 BP, indicating
perennial water with either greater precipitation or less evaporation.
Deposit 4 was an aeolian sand that was likely contemporary with the
first advance of the Nafud Desert, thereby marking a period of greatly
increased aridity. Two sites with Levallois elements (201-25a,b;
201-26a) were identified in the Jubbah Basin (Figure 4). Site 201-26a
was located close to a mountainous summit, the site representing a
quarry based on the exploitation of the ferruginous sandstone outcrop.
Site 201-25a was on a lower sandstone platform and contained ferruginous
sandstone and local quartzite artefacts. Geomorphic observations
indicated the presence of a lake basin surrounded by dunes. A scatter of
"Levalloiso-Mousterian" artefacts was found on the weathered
surface of 201-25b (Deposit 5). Since Deposit 5 was radiocarbon dated to
25 630 BP, the investigators assumed that the artefacts were washed onto
the surface.
[FIGURES 3-4 OMITTED]
The temporal sequence outlined for the Jubbah Basin in the north
appears to overlap with the deposits identified in 24-metre long
profiles established for Lake Mundafan in the Rub' al-Khali
(McClure 1976, 1978). The fluctuations of wet and arid phases documented
in the Jubbah and Lake Mundafan sedimentary sequences are likely tied to
climatic changes occurring during the Quaternary. Although
archaeological research has not been well-integrated with
palaeoenvironmental studies, it is likely that climatic cycles heavily
influenced the occupation history of Arabia. Study of such relationships
is of profound importance as environmental stress and aridity have been
considered to be important factors in conditioning hominin biogeography,
dispersals, and adaptations (Lahr & Foley 1998; Walter et al. 2000).
Although only known from preliminary information, archaeological
surveys in Arabia have indicated variations in site density and
distribution. Survey results in certain parts of Arabia indicated that
Middle Palaeolithic sites were not present or present in small numbers.
In other cases, large numbers of sites were reported, such as the
presence of 31 sites in a survey in northern Saudi Arabia (Gilmore et
al. 1982). Indicative of activity variations, artefact densities ranged
from small numbers of finds to thousands of artefacts per site, and in
one survey, surface distributions were reported to range from 375 to 12
500 [m.sup.2] (e.g. Whalen & Pease 1992).
Technology and assemblage characteristics
Middle Palaeolithic core reduction methods of Arabia have been
described as plain flake, discoidal, Levallois, and blade. Resultant
artefact types include distinctive debitage and retouched tools (Figure
5). Researchers working in Arabia have repeatedly noted that
"Levallois" core technology" was not as well represented
as Levantine Mousterian industries, thereby indicative of technological
variability. The meaning of typo-technological systematics have been
interpreted as signatures of population movements and responses to
climatic change (e.g. Bar-Yosef 1994) or the product of a host of
adaptive options, such as raw material types and distributions, mobility
strategies, and site duration (e.g. Clark 2000).
[FIGURE 5 OMITTED]
The traditional use of the term 'Mousterian' to describe
Arabian assemblages implies that this region shares general stylistic
affinities with those in Europe, western and central Asia, and North
Africa. Yet, many assemblages classified as Mousterian in Arabia were
noted to contain relatively low frequencies of Levallois cores or
elements, apparently departing from particular areas of Europe and the
Levant which show high frequencies of Levallois technology. Three
particular variants of the Arabian Middle Palaeolithic were identified,
classed as "Mousterian of Acheulean Tradition" (MAT),
"pebble Mousterian", and "Aterian". Tool indices
from Arabian sites were argued to correspond with MAT Type A assemblages
found in other parts of Eurasia (Whalen etal. 1981: table 3). Another
possibility is that small bifaces in prepared core assemblages may
represent transitional assemblages across the Upper Acheulean and early
Middle Palaeolithic (e.g. Bar-Yosef 1998; Petraglia et al. 2003).
Although the pebble Mousterian was noted to be a particular
technological variant (Parr et al. 1978), it is more likely that the
resulting tool forms are related to clast size and form, such as found
in the Pontinian (Kuhn 1995).
Of significance to understanding inter-regional trends, a single
site on the south-west edge of the Rub' al-Khali was classified as
Aterian (McClure 1994). A total of 300 tools was recovered across an
area measuring 100 [m.sup.2.] The assemblage was characterised by large,
unifacially worked, predominately tanged points and scrapers, as well as
miscellaneous types (small bifacial foliates, scrapers, knives, awls,
denticulates) that were tanged, and non-tanged (Figure 6). Tangs usually
displayed retouch on the dorsal flake surface, although ventral working
was occasionally applied. The tools were manufactured from the same type
of tan-coloured flint, indicating reduction from a common source. The
site was inferred to be associated with a well-watered grassland
environment. The identification of an Aterian assemblage on the Arabian
Peninsula represents a provocative case for potential geographic
connections with populations in North Africa. The single Aterian example
potentially extends the eastward range of this point technology outside
of Africa. The point technology may also indicate similarities in
adaptations over a broad area, signalling a shift to new forms of
hunting tactics during periods of improved environmental conditions.
[FIGURE 6 OMITTED]
Apart from the three identified variants of the Middle
Palaeolithic, the overall impression is that most stone tool assemblages
of Arabia show flake and undifferentiated industries. Although broadly
similar in core and flake technology, the Arabian "Mousterian"
assemblages are not clear equivalents with the Levantine and Zagros
Mousterian. In part, this may have to do with the fact that the
character of the Arabian assemblages have not been adequately
distinguished through systematic typological studies in
chrono-stratigraphic contexts. Certain researchers have hypothesised
that the Arabian Peninsula followed its own technological development,
incorporating a different cultural stylistic tradition and representing
a specific adaptation to particular environmental conditions (Whalen et
al. 1981). Although technological and geographic heterogeneity is an
intriguing idea, the degree to which the Arabian industries share
technological features with outlying regions must remain an open
question.
The presence of flake and blade elements in the Middle Palaeolithic
of the Arabian Peninsula is of interest to typological systematics and
adaptive behaviours. The presence of a blade component in Arabian Middle
Palaeolithic assemblages is consistent with findings from Africa and the
Levant, indicating that these technologies were used much earlier than
previously surmised (e.g. Bar-Yosef & Kuhn 1999). The predominance
of flake-based technologies over blade bearing industries later in time
led certain researchers to remark that Mousterian toolkits may have a
longer survivorship in the Arabian Peninsula (Zarins et al. 1980;
Alsharekh 1995). In this view, the late occurrence of Middle
Palaeolithic technology overshadows the rarer blade bearing industries
of the Upper Palaeolithic. The Arabian situation may show a different
Middle to Upper Palaeolithic trajectory from other regions, supporting
the notion that a uniform and directional change from flake-to
blade-based industries should not be expected.
With respect to procurement, Middle Palaeolithic assemblages
consist of a wide range of raw materials derived from primary bedrock
and secondary gravel sources. The favoured raw material was quartzite
and flint, although other raw materials were heavily used in certain
areas, including rhyolite, andesite, and basalt. Many Middle
Palaeolithic sites are either directly associated with raw material
outcrops or in close spatial proximity to them. Large "factory
sites" were found along chert and ferruginous sandstone outcrops,
the collections from such sites consisting of cores, flakes, and
retouched flake tools (Figure 7). It is not known how far materials were
transported, but most appear to be confined within basins and valleys,
and not at great distances from sources, paralleling observations in
other regions.
[FIGURE 7 OMITTED]
Hominin dispersals
Currently available evidence shows that the Arabian Peninsula was
occupied by hominins employing Middle Palaeolithic technologies, thereby
providing information about hominin colonisation and dispersal processes
(Figure 8). The presence of Middle Palaeolithic sites in the peninsula
complements the identification of Lower Palaeolithic sites, most of
which are Acheulean (Petraglia 2003). It is uncertain, however, whether
early Middle Palaeolithic technology developed from the Late Acheulean,
as was found in other regions outside of Africa (e.g. Petraglia et al.
2003; White & Ashton 2003), or whether early Middle Palaeolithic
technologies of Arabia were introduced as part of hominin dispersals.
While there are uncertainties about the evolution of Middle Palaeolithic
technologies, general observations indicate that there are gross
similarities between Middle Palaeolithic industries in Africa, Arabia,
and South Asia based on the presence of prepared core industries
(Petraglia et al 2003). This geographic observation for prepared core
industries stands in contrast to contemporaneous sites in eastern Asia,
which are characterised by pebble core technology similar to reduction
techniques used in the Lower Palaeolithic (e.g. Gao & Norton 2002).
It is possible that inter-regional differences in assemblages reflect
divergences in resource bases, hominin biogeography, and adaptations.
[FIGURE 8 OMITTED]
With respect to the hominins using Arabian technology, a few
possibilities exist, as Middle Palaeolithic industries are thought to be
associated with at least two, if not three species, including H. helmei,
H. neanderthaknsis, and H. sapiens (e.g. Klein 1999; McBrearty &
Brooks 2000; Lahr & Foley 2001). Although no human fossils have been
recovered in Arabia, it is likely that Middle Palaeolithic assemblages
were produced by early and later modern humans, potentially including H.
helmei and H. sapiens. Another more remote possibility, in consideration
of the close geographic proximity of the Levantine Mousterian and the
Zagros Mousterian, is that the Arabian industries represent the southern
extension of the Neanderthals.
The Out of Africa model for modern human origins and the eastward
migration of hominin populations has been supported by fossil and
genetic studies (e.g. Lahr & Foley 1998; Quintana-Murci et al. 1999;
Stringer 2002). Although the Arabian assemblages share generic
technological affinity with Middle Palaeolithic industries in Africa and
the Levant, it is premature at this stage to infer population
connections without detailed inter-assemblage comparisons. Yet, mapping
of the Arabian sites does provide information that is relevant for
examining dispersal processes. Site distributions support an argument
that more than one route was taken by populations emerging out of
Africa. One route may be from northern Africa into the Levant,
simultaneously or subsequently leading to the occupation of Arabia. Once
populations were present in Arabia, they may have underwent contraction
during periods of aridity. Coastal migrations are suggested by the
presence of archaeological sites along the Red Sea and the Arabian Sea.
Populations may have crossed the Bab al Mandab during lowstands or they
may have employed boat technology to cross the Strait. Particularly
intriguing is the presence of 25 Middle Palaeolithic sites near the Bab
al Mandab (Whalen & Pease 1992). The Red Sea margin sites may relate
to northward expansion of human populations. A second possibility is
that the Red Sea sites may represent the spread of populations moving
southwards. Evidence for sites along the Arabian Sea margin may
represent the dispersal of populations expanding eastwards, moving along
shorelines themselves, or possibly employing boats, as inferred for the
colonisation of Australia by 60 000 BP (Stringer 2000). The use of
coastal margins in Arabia may relate to evidence for marine adaptations
in East Africa by 125 000 BP (Walter et al. 2000). Although connections
between populations are difficult to prove from technological evidence
alone, the presence of an Aterian site in Arabia provides a potential
link, or at least an adaptive similarity with populations in northern
Africa.
Conclusions
Archaeological research in Arabia has indisputably confirmed the
presence of Middle Palaeolithic sites in a number of areas of the
peninsula. Despite the importance of the Arabian Middle Palaeolithic to
palaeoanthropological models, there has been no systematic attempt to
examine how the Arabian industries compare with the Mousterian of West
Asia and the MSA of Africa. Of course, chronometric dating and in-depth
technological studies are critical for ascertaining how the Arabian
record fits into inter-regional dispersal processes and evolutionary
sequences. Based on the preliminary evidence, the Arabian Peninsula
likely acted as a key route for dispersing populations given the
identification of a number of sites in the region and the presence of
sites along the Red Sea and Arabian Sea margins.
Given the marked environmental changes in the Arabian Peninsula and
the presence of archaeological sites in different ecological settings,
it is clear that the regional record can contribute valuable information
about hominin responses to changing conditions. The record shows that
Middle Palaeolithic occupations were associated with wet environments
and favourable resources. A large unanswered question is how hominins
coped with periods of environmental degradation, including increased
aridity. The persistence and spatial distribution of Middle Palaeolithic
assemblages in Arabia suggests that hominins may have had some cultural
abilities to adjust to ecological challenges. Without further detailed
analysis of palaeoecological and archaeological settings, it is
difficult to ascertain the degree to which hominins adjusted to
inhospitable conditions and whether populations contracted in size or
became extinct.
Based on a review of the currently available literature, it is
clear that the Middle Palaeolithic of the Arabian Peninsula offers
unique opportunities to understand Pleistocene dispersals, adaptations,
and behaviour. Current shortcomings in palaeoanthropological research
will hopefully be rectified in the near future based upon new
initiatives being undertaken jointly between the Department of
Antiquities and Museums (Saudi Arabia) and UK universities. Recent
support from the Natural Environment Research Council (NERC) to the
Leverhulme Centre for Human Evolutionary Studies, University of
Cambridge, provides the opportunity to develop collaborative efforts
with the Saudi National Museum and King Saud University.
Interdisciplinary teams are being formed to examine the chronology of
human occupation, the impact of palaeoenvironmental changes on
populations, and the dispersal of modern humans outside of Africa.
Acknowledgements
A Fulbright grant to Petraglia stimulated our collaborative
research efforts. Numerous colleagues in Saudi Arabia have supported and
facilitated our research, especially Prof. Saad Abdul-Aziz Al-Rashid,
All S. al-Moghanam, Jamal-Saraj, and Saad M. Al-Mashari. Petraglia
wishes to acknowledge the support and assistance of Lora Berg, Peter
Lenthall, Paul Taylor, Peter Vincent, and Norman Whalen. We are grateful
to Larry Barham, Nicole Boivin, Martin Carver, Marta Lahr, Preston
Miracle and Yosuf El-Amin for comments on a draft of this article.
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Michael D. Petraglia (1)& Abdullah Alsharekh (2)
(1) Leverhulme Centre for Human Evolutionary Studies, University of
Cambridge, Downing Street, Cambridge 6732 3DZ, England (Email:
[email protected])
(2) Department of Archaeology and Museology, King Saud University
170. Box 2456, Riyadh 11451, Kingdom of Saudi Arabia